The centerlock "Flushlock" system converts the lug studs into drive pins. Then those pins engage the back side of the wheel, in holes that are drilled to match the pin location. Just like a normal centerlock setup. That large spline nut actually holds the wheel in place.
That particular set above (silver/red) is actually for a Gen 2 Viper GTS, local customer.
Here is how it looks on the Gen 5 Viper.
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